皖北地区寒武系芙蓉统炒米店组中上部的索克虫类三叶虫_雷倩萍.pdf
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1、 收稿日期:2021-12-11;改回日期:2022-06-17;录用日期:2022-08-22*国家自然科学基金(41802004)、中国地质调查局项目(DD20190009)和现代古生物学和地层学国家重点实验室项目(20191101)联合支持。*通讯作者:雷倩萍,副研究馆员,e-mail: 古生物学报,61(4):568589(2022年12月)Acta Palaeontologica Sinica,61(4):568589(December 2022)研究论文 DOI:10.19800/ki.aps.2021071 皖北地区寒武系芙蓉统炒米店组中上部的 索克虫类三叶虫*雷倩萍1*彭善池2
2、,3 1 常州博物馆学术研究部,江苏常州 213022;2 中国科学院南京地质古生物研究所,南京 210008;3 现代古生物学和地层学国家重点实验室,南京 210008 提要 在安徽省萧县凤凰山剖面炒米店组中上部(江山阶上部到第十阶)岩层中采得丰富的索克虫科三叶虫标本。共计4属6种(含2新种),包括Eosaukia bella(Walcott,1906)、E.anhuiensis sp.nov.、Lophosaukia orientalis(Kobayashi,1933)、Prosaukia campe(Walcott,1905)和P.xiaoxianensis sp.nov.以及Liche
3、ngia onigawara Kobayashi,1942。在对属征进行修订与讨论的基础上,通过对比这些属的模式种及其他种,对4个旧种进行了再研究和种征厘定,最后对这6个种的谱系关系开展了简要讨论。关键词 索克虫科 三叶虫 芙蓉统 寒武系 皖北地区 中文引用 雷倩萍,彭善池,2022.皖北地区寒武系芙蓉统炒米店组中上部索克虫科三叶虫.古生物学报,61(4):568589.DOI:10.19800/ki.aps.2021071 英文引用 Lei Qian-ping,Peng Shan-chi,2022.Saukiid trilobites from the middleupper Chaumit
4、ien Formation(Fu-rongian,Cambrian)in northern Anhui.Acta Palaeontologica Sinica,61(4):568589.DOI:10.19800/ki.aps.2021071 Saukiid trilobites from the middleupper Chaumitien Formation(Furongian,Cambrian)in northern Anhui LEI Qian-ping1,PENG Shan-chi2,3 1 Academic Department of Changzhou Museum,Changzh
5、ou 213022,Jiangsu,China;2 Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences,Nanjing 210008,China;3 State Key Laboratory of Palaeobiology and Stratigraphy,Nanjing 210008,China Abstract Hundreds of specimens of saukiid trilobites were collected from the middleupper Chaumitien
6、Formation(upper Jiangshanian through Stage 10,Cambrian)at Fenghuangshan Hill in northern Anhui,China.Four genera and six species(including two new species)are described herein:Eosaukia bella(Walcott,1906),E.anhuiensis sp.nov.,Lo-phosaukia orientalis(Kobayashi,1933),Prosaukia campe(Walcott,1905)and P
7、.xiaoxianensis sp.nov.,Lichengia onigawara Kobayashi,1942.After revising and discussing the generic diagnoses,we reexamine and revise the specific diagnoses of the four previously-published species by comparing their type species with others,and briefly discuss the evolutionary relationships among a
8、ll six species.Key words Saukiidae,trilobite,Furongian,Cambrian,northern Anhui 4期 雷倩萍、彭善池:皖北寒武三叶虫;Lei and Peng:Saukiid trilobites from Cambrian in Anhui 569 SYSTEMATIC PALEONTOLOGY All specimens described in this paper are deposited at the Changzhou Museum(CZM)and Nanjing Institute of Geology and Pa
9、laeontology,Chinese Academy of Sciences(NIGP).Order Asaphida Salter,1864 Superfamily Dikelocephaloidea Miller,1889 Family Saukiidae Ulrich et Resser,1930 Genus Eosaukia Lu,1954 Eosaukia anhuiensis sp.nov.urn:lsid:zoobank.org:act:EC4770D0-BC7C-418E-A4CF-6AB8328FF4C1(Figs.5,11)Etymology After Anhui Pr
10、ovince,where the specimens are collected.Holotype Cranidium(CZMF000143;Fig.5-11)collected from the Chaumitien Formation at Fenghuangshan,Xiaoxian,Anhui Province.Materials Twenty-eight cranidia,12 libregenae,and seven pygedia collected from the interval of WXN-7h to WXN-3k.Diagnosis Cranidium trapezo
11、id,slightly convex.Glabella long,trapezoid,narrowed forward.Four gla-bellar furrows:S1 traverse,curved rearward and deep-ened laterally;S2 short and deep;S3,S4 short and shal-low;occipital ring medium-sized,occipital spine occa-sionally present.Palpebral lobe medium-sized.Anterior area wide(sag.);la
12、teral anterior border furrow of cra-nidium diagonal and shallow.Librigena convex.Lateral border narrow.Genal spines long,hook-like and strongly curved inward.Pygedium triangular,axis wide(tr.),with three axial rings and a terminal piece;pleural furrows and interpleural furrows indistinct.Exoskeleton
13、 with granules.Description Cranidium trapezoid,slightly con-vex,length about four-fifths of width.Glabella sub-trapezoid,slightly tapered forward,expanded rear-ward posterior to the mid-length,width about two-thirds of length(sag.).Four pairs of glabellar furrows,S1 traverse,curved rearward and deep
14、ened laterally;S2 short(tr.),about one-third of glabellar width,curved rearward;S3 short and shallow,not connected with axial furrows;S4 short and shallow,slightly directed forward.Occipital ring with uniform width,curved rearward;oc-cipital furrow deep and broad,curved rearward;occipi-tal spine occ
15、asionally present(Figs.5-8,5-11,5-12).Anterior area wide(sag.),narrowing laterally,and forming an obtuse angled anterior margin,about two-fifths as long as glabellar width;lateral anterior border furrow of cranidium diagonal,wide and shallow.Axial furrows deep and wide.Palpebral lobe moderately size
16、d,extending between S2 and half-length of L1;pal-pebral furrow deep.Palpebral area about one-third as wide as glabella.Facial suture opithoparian:anterior branch of facial suture subparallel,curving axially close to cephalic anterior margin;posterior branch of facial suture gently extending outward,
17、about 20 curved pos-terior margin of cranidium.Posterolateral projection short(exs.),with width about glabellar length.Cra-nidium with granules.Librigena gently convex.Librigenal field sharply sloping to the lateral border;eye socle furrow medium width and deep;lateral border narrow(Fig.5-18)and con
18、vex,extending rearward and outward,then gradually curving inward to form a hook-like genal spine;lateral border furrow wide and shallow,forming a depression at the genal angle.Librigena surface with granules.Pygidium convex,semi-oval,width two times of length.Axis strongly convex,two times as long a
19、s width,narrowing rearward to the posterior border with three axial rings and a terminal piece.Pleural furrow deep and gently extending rearward and outward;inter-pleural furrow shallow.Pygidial border narrow.Comparison and discussion The new species differs from the type species in having a larger
20、and wid-er eye lobe,and a narrower(sag.)and longer pygidium;Eosaukia walcotti(Mansuy,1915)from Vietnam differs from the new species in having a narrower(sag.)anteri-or area,a librigena without strongly hooked spine;Eo-saukia bella(Walcott,1906)differs from the new spe-cies in having a narrower(sag.)
21、and more traverse ante-rior area,a more convex exoskeleton,a relatively shorter genal spine which is not strongly hooked,and a wider pygidial pleural field;Eosaukia transita(Lu and Zhou,1990)differs from the new species in having a more traverse anterior area,a narrower(sag.)occipital ring,and no oc
22、cipital spine;Eosaukia tangcunensis(Qian,1983)differs from the new species in having a wider(sag.)anterior area,a more rounded glabella at the front,and a narrower fixigena;Eosaukia combinata(Peng,1992)differs from the new species in having a paral-lel-sided glabella and a greater curved eye lobe.Th
23、e occipital spine may be present or absent dur-ing the various stages of the ontogeny(Sherglod,1991).The occipital spine may also be caused by dimorphism or taphonomy,such as the case that some of our speci-mens have occipital spines(Figs.5.8,5.11,5.12),while some have not.Occurrence Middleupper Cha
24、umitien Formation,Cambrian,Furongian,northern Anhui.Prosaukia xiaoxianensis sp.nov.urn:lsid:zoobank.org:act:1F502F4C-C9F2-4C88-A9C7-A20C9B1DA971(Figs.9,11)570 古 生 物 学 报 61 卷?1935 Tellerina chinhsiensis Sun,p.48,pl.6,fig.19,not figs.1518.?1965 Ptychaspis shansiensis Sun,Lu et al.,p.426,pl.82,fig.10,n
25、ot figs.79,11,12.?1965 Tellerina chinhsiensis Sun,Lu et al.,p.455,pl.89,fig.11,not figs.910.?1965 Tellerina sulcatifera Endo,Lu et al.,p.456,pl.89,figs.1516,not figs.1314,17.Etymology After Xiaoxian,Anhui Province,where the specimens are collected.Holotype Cranidium(CZMF000180;Fig.9-5)collected from
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